An answer for "Why does the small h letter in Garamond italic bend inward?" shows the following image:
The middle row of this image (marked Han 86R, Rome 1468) has the capital N inverted compared to the others, which range from 1465 to 1470. Note that the top serifs are consistently longer on the left, and the bottom serifs are consistently longer on the left. This shows that it was a deliberate design choice, not an accidentally mirrored photo.
Original:
Flipped horizontally:
Flipped vertically:
As you can see, flipping the image produces a weird-looking N, because the serifs are long in the wrong direction.
Why was the N inverted in just this typeface?
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